Plants: Monoecious. Ovulate cones: Compound; cone bracts spirally arranged, flattened, tongueshaped, free from scale, with a single vascular strand; ovuliferous scales flattened; ovules 2, inverted, proximal, fused to ovuliferous scale, with a single vascular strand dividing up to 20 times, micropyle laterally directed; seeds typically winged, rarely unwinged (e.g. Pinus pinea), wing descended from ovuliferous scale. Pollen cones: Simple, sometimes clustered on short shoots originating from a commen bud; sporangiophores several, hyposporangiate, always with 2 sporangia; pollen mostly with two distinct air-bladdes. Leaves: Simple, arranged either along long-shoots (persistent), or on long- and short-shoots (deciduous or persistent), or only on short-shoots (persistent) with two vascular strands surrounded by a common bundle sheath.
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