Features of genus those of family. Endarch stele with uni-biseriate bordered pits on secondary tracheids, and rays 1–2 cells wide, resin rodlets in ground tissues. Leaves amphistomatic; adaxial surface with two long stomatal bands each 4–8 stomatal complexes wide with some shared subsidiary cells, diminishing to single rows or scattered stomata near tip of small leaves; abaxial surface with two shorter basal bands or scattered basal stomata. Stomata monocyclic with unipapillate subsidiary cells.
Scratchpads developed and conceived by (alphabetical):
Ed Baker,
Katherine Bouton
Alice Heaton
Dimitris Koureas,
Laurence Livermore,
Dave Roberts,
Simon Rycroft,
Ben Scott,
Vince Smith